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> New anatomic studies review the ability to walk upright and  climb trees:
> 
> https://phys.org/news/2019-02-upright-human-ancestors.html
> 
> "Scott W. Simpson, Ph.D., led an analysis of a 4.5 million-year-old fragmentary female skeleton of the human ancestor Ardipithecus ramidus that was discovered in the Gona Project study area in the Afar Regional State of Ethiopia.
> 
> "The newly analyzed fossils document a greater, but far from perfect, adaptation to bipedalism in the Ar. ramidus ankle and hallux (big toe) than previously recognized. "Our research shows that while Ardipithecus was a lousy biped, she was somewhat better than we thought before," said Simpson.
> 
> ***
> 
> "The new analysis, published in the Journal of Human Evolution, thus points to a diversity of adaptations during the transition to how modern humans walk today. "The fact that Ardipithecus could both walk upright, albeit imperfectly, and scurry in trees marks it out as a pivotal transitional figure in our human lineage," said Simpson.
> 
> "Key to the adaptation of bipedality are changes in the lower limbs. For example, unlike monkeys and apes, the human big toe is parallel with the other toes, allowing the foot to function as a propulsive lever when walking. While Ardipithecus had an offset grasping big toe useful for climbing in trees, Simpson's analysis shows that it also used its big toe to help propel it forward, demonstrating a mixed, transitional adaptation to terrestrial bipedalism.
> 
> "Specifically, Simpson looked at the area of the joints between the arch of the foot and the big toe, enabling him to reconstruct the range of motion of the foot. While joint cartilage no longer remains for the Ardipithecus fossil, the surface of the bone has a characteristic texture which shows that it had once been covered by cartilage. "This evidence for cartilage shows that the big toe was used in a more human-like manner to push off," said Simpson. "It is a foot in transition, one that shows primitive, tree-climbing physical characteristics but one that also features a more human-like use of the foot for upright walking." Additionally, when chimpanzees stand, their knees are "outside" the ankle, i.e., they are bow-legged. When humans stand, the knees are directly above the ankle—which Simpson found was also true for the Ardipithecus fossil.
> 
> Comment: this an obvious transitional fossil, but full blown speciation requiring design. I don't believe the cells of the common ancestor of chimps and humans could conceive of how to design a foot and spine and pelvis for  bipdal movement.